Publications *student co-author
32. Krug, P.J., Vendetti, J.E., Retana*, J., Rodriguez*, A., Hirano, Y., and Trowbridge, C.D. 2013. DNA barcoding of mitochondrial and
nuclear loci reveals rampant cryptic speciation among sea slugs studied for drug discovery, plastid symbiosis and biological control.
Molecular Phylogenetics and Evolution, accepted pending revision.
31. Romero*, M.R., Phuong*, M., Bishop, C.D., and Krug, P.J. 2013. Differential function of nitric oxide signaling during habitat
choice by two larval morphs of the sea slug Alderia willowi: mechanistic insight into evolutionary transitions in dispersal strategies.
Journal of Experimental Biology, in press.
30. Marshall, D. J., Krug, P.J., Kupriyanova, E., Byrne, M., and R. B. Emlet. 2012. The biogeography of marine invertebrate life histories.
Annual Reviews of Ecology, Evolution and Systematics, 43: 97-114. (doi: 10.1146/annurev-ecolsys-102710-145004)
29. Krug, P.J., Gordon*, D., and M. Romero*. 2012. Seasonal polyphenism in larval type: Rearing environment influences the development
mode expressed by adults in the sea slug Alderia willowi. Integrative and Comparative Biology, 52: 161-172. (doi: 10.1093/icb/ics059)
28. Vendetti, J.E., Trowbridge, C.D., and P. J. Krug. 2012. Poecilogony and population genetic structure in Elysia pusilla
(Heterobranchia: Sacoglossa), and reproductive data for five sacoglossans that express dimorphisms in larval development.
Integrative and Comparative Biology, 52: 138-150. (doi: 10.1093/icb/ics077) PDF Table_S1
27. Krug, P.J., Asif*, J.H., Baeza*, I., Morley, M., Blom, W. and T. Gosliner. 2012. Molecular identification of two species of the carnivorous
sea slug Philine, invaders of the U.S. west coast. Biological Invasions, 14: 2447-2459. (doi: 10.1007/s10530-012-0242-9) PDF Table_S1
26. Asif*, J.H. and P.J. Krug. 2012. Lineage distribution and barriers to gene flow among populations of a globally invasive marine mollusc.
Biological Invasions, 14: 1431-1444. (doi: 10.1007/s10530-011-0169-6) PDF supplemental_material
25. Krug, P.J., Händeler, K., and J. Vendetti. 2011. Genes, morphology, development and photosynthetic ability support the resurrection of
Elysia cornigera (Heterobranchia: Plakobranchoidea) as distinct from the solar-powered sea slug E. timida.
Invertebrate Systematics, 25: 477–489. (doi: 10.1071/IS11026) PDF Table_S1
24. Krug, P. J. 2011. Patterns of speciation in marine gastropods: A review of the phylogenetic evidence for localized radiations in the sea.
American Malacological Bulletin, 29: 169-186. (doi: 10.4003/006.029.0210) PDF
23. Händeler, K., Grzymbowski, Y.P., Krug, P.J. and H. Wägele. 2009. Functional chloroplasts in metazoan cells – a unique evolutionary
strategy in animal life. Frontiers in Zoology, 6: 28 http://www.frontiersinzoology.com/content/6/1/28
22. Allen*, R.M., Krug, P.J., and D.J. Marshall. 2009. Larval size is determined by extra-embryonic provisioning but not egg size in a marine
gastropod. Marine Ecology Progress Series, 389: 127-137. PDF
21. Krug, P.J. 2009. Not my "type": Bet-hedging and dispersal dimorphisms in opisthobranch life histories. Biological Bulletin, 216:
355-372. PDF
20. *Smolenksy, N., *Romero, M. R., and P.J. Krug. 2009. Evidence for costs of mating and self-fertilization in a simultaneous hermaphrodite
with hypodermic insemination, the opisthobranch Alderia willowi. Biological Bulletin, 216: 188-199. PDF
19. Krug, P.J., Riffell, J.A., and R. K. Zimmer. 2009. Endogenous signaling pathways and chemical communication between sperm and egg.
Journal of Experimental Biology, 212: 1092-1100. PDF
18. Krug, P.J., Morley, M.S., *Asif, J.H., Hellyar, L.L. and W. M. Blom. 2008. Molecular confirmation of species status for the rare
cephalaspidean Melanochlamys lorrainae (Rudman, 1968), and comparison with its sister species M. cylindrica Cheeseman, 1881.
Journal of Molluscan Studies, 74: 267-276. PDF
17. Krug, P.J. 2007. Poecilogony and larval ecology in the gastropod genus Alderia. American Malacological Bulletin 23: 99-111. PDF
16. Krug, P.J., *Ellingson, R.A., Burton, R.A. and Á. Valdés. 2007. A new poecilogonous species of sea slug (Opisthobranchia: Sacoglossa)
from California: Comparison with the planktotrophic congener Alderia modesta (Lovén, 1844). Journal of Molluscan Studies 73: 29-38. PDF
15. *Ellingson, R.A. and P.J. Krug. 2006. Evolution of poecilogony from planktotrophy: Cryptic speciation, phylogeography and larval
development in the gastropod genus Alderia. Evolution 60: 2293-2310. PDF
14. Zimmer, R.K., Schar, D.W., Ferrer, R.P., Krug, P.J., Katz, L.B., and W.C. Michel. 2006. The scent of danger: Tetrodotoxin (TTX)
as an olfactory cue of predation risk. Ecological Monographs 76: 585-600. PDF
13. *Botello, G. and P.J. Krug. 2006. Desperate larvae revisited: Age, energy and experience affect sensitivity to settlement cues in larvae
of the gastropod Alderia sp. Marine Ecology Progress Series 312: 149-159. PDF
12. Krug, P.J. 2006. “Defense of benthic invertebrates against surface colonization by larvae: A chemical arms race.” Pp. 1-53 in:
Clare A. and Fusetani N. (eds) Marine Molecular Biotechnology, vol 2. Springer, Berlin. PDF
11. Krug, P.J. and and R.K. Zimmer. 2004. Developmental dimorphism: Consequences for larval behavior and dispersal potential in a
marine gastropod. Biological Bulletin 207: 233-246. PDF
10. Riffell, J.A., Krug, P.J., and R.K. Zimmer. 2004. The ecological and evolutionary consequences of sperm chemoattraction.
Proceedings of the National Academy of Sciences 101: 4501-4506. PDF
9. Riffell, J.A., Krug, P.J., and R.K. Zimmer. 2002. Fertilization in the sea: The chemical identity of an abalone sperm attractant.
Journal of Experimental Biology 205: 1439-1450. PDF
8. Krug, P.J. 2001. Bet-hedging dispersal strategy of a specialist marine herbivore: A settlement dimorphism among sibling larvae
of Alderia modesta. Marine Ecology Progress Series 213: 177-192. PDF
7. Krug, P.J. and and R.K. Zimmer. 2000. Developmental dimorphism and expression of chemosensory-mediated behavior: Habitat
selection by a specialist marine herbivore. Journal of Experimental Biology 203: 1741-1754. PDF
6. Krug, P.J. and and R.K. Zimmer. 2000. Larval settlement: chemical markers for tracing production, transport, and distribution of
a waterborne cue. Marine Ecology Progress Series 207: 283-296. PDF
5. Krug, P.J., and A.E. Manzi, 1999. Waterborne and surface-associated carbohydrates as settlement cues for larvae of the specialist
marine herbivore Alderia modesta. Biological Bulletin 197: 94-103. PDF
4. Krug, P.J., 1998. Poecilogony in an estuarine opisthobranch: Planktotrophy, lecithotrophy, and mixed clutches in a population of
the estuarine opisthobranch Alderia modesta. Marine Biology 132: 483-494. PDF
3. Graziani, E.I., Andersen, R.J., Krug, P.J., and D.J. Faulkner. 1996. Stable isotope incorporation incorporation evidence for the
de novo biosynthesis of terpenoic acid glycerides by dorid nudibranchs. Tetrahedron 52: 6869-6878. PDF
2. Krug, P.J., Boyd, K.G., and D.J. Faulkner. 1995. Isolation and synthesis of Tanyolides A and B, metabolites of the nudibranch
Sclerodoris tanya. Tetrahedron 51: 11063-11074.
1. Krug, P.J., Gileski, A.Z., Code, R., Torjussen, A., and M.B. Schmid. 1994. Endpoint bias in large Tn-10 catalyzed inversions in
Salmonella typhimurium. Genetics 136: 747-756. PDF